DMMC Course: UNRAVELLING CHROMATIN & THE ROLE OF EPIGENETICS IN DISEASE
1030-1100 Tuesday 25 April 2006. UCD Conway Institute Lecture Theatre
DNA
methylation
Dr
Colum Walsh (School of Biomedical Sciences, University of Ulster)
Methyl groups are small chemical tags which are added to the cystosine in eukaryotic DNA after replication. This initially occurs in the early stages of development due to the action of one class of enzymes, called de novo methyltransferases. Later in life these disappear, but methylation is faithfully maintained by the action of the maintenance enzyme DNMT1. In this lecture, I will talk about what we know methylation is needed for, and its importance in development and in cancer. The role of the methyltransferase enzymes and the proteins which bind methylated DNA to interpret the signal will also be discussed.
Resources
References
Bestor, T. H. (2000). The DNA methyltransferases of mammals. Hum. Mol. Genet. 9, 2395-2402. PubMed Entry
Bird, A. (2002). DNA methylation patterns and epigenetic memory. Genes Dev. 16, 6-21. PubMed Entry
Eden, A., Gaudet, F., Waghmare, A. & Jaenisch, R. (2003). Chromosomal instability and tumors promoted by DNA hypomethylation. Science 300, 455. PubMed Entry
Ting, A. H. et al. (2004). CpG island hypermethylation is maintained in human colorectal cancer cells after RNAi-mediated depletion of DNMT1. Nat. Genet. 36, 582-584. PubMed Entry
Xu, G. L. et al. (1999). Chromosome instability and immunodeficiency syndrome caused by mutations in a DNA methyltransferase gene. Nature 402, 187-191. PubMed Entry
Walsh, C. P., Chaillet, J. R. & Bestor, T. H. (1998). Transcription of IAP endogenous retroviruses is constrained by cytosine methylation. Nat. Genet. 20, 116-117. PubMed Entry
Web Links
The Catalogue of Imprinted Genes and Parent-of-origin Effects in Humans and Animals: http://igc.otago.ac.nz/home.html
geneimprint.com: http://www.geneimprint.com/
The
Epigenome Network of Excellence (NoE): http://www.epigenome-noe.net/